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Unit Evolution Gallery

🔄 Two Evolution Systems • ⭐ E1-E6 Progression • 💎 Material Requirements • 🏆 Endgame Power

Explore the two evolution systems in Universal Tower Defense. Base Evolution transforms units with materials into powerful new forms. Etherealization (E1-E6) progressively enhances units through stage-based upgrades.

🔄 Base Evolution⭐ E1-E6 Ethereal Stages📦 Material System✅ Updated Dec 2025
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🔄

What is Base Evolution?

Base Evolution is a one-time transformation system that fundamentally changes a unit's form, abilities, and stats. By collecting specific evolution materials and reaching Level 50, you can evolve select units into significantly more powerful versions with new abilities and enhanced base stats.

🔥Transform Unit Form
💪Enhanced Base Stats
✨New Abilities & Passives
📦Material Requirements
⭐

What is Etherealization?

Etherealization is a progressive enhancement system in Universal Tower Defense. By collecting duplicate units or specific evolution materials, you can advance your units through six evolution stages (E1-E6). Each stage unlocks powerful passive abilities and provides significant stat bonuses, transforming units into endgame powerhouses.

🎯Unlock Unique Passives
📈Massive Stat Increases
💎Material-Based System
🏆Endgame Essential
🔍

Showing 15 of 15 evolution paths

Evolution Path

Base Form

Spade

DPS:420
Range:20
Rarity:Mythic
→
💰25,000
⭐Level 50+
Evolved Form

Spade (Unleashed)

DPS:1450+245%
Range:35
Rarity:Mythical

📦 Required Materials

Flame Essence×10
Fire Stone×5

✨ New Ability

Enkai: Hibashira

Creates a massive pillar of fire that deals 3x base damage and applies intense burn.

Cooldown: 40s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Rule

DPS:380
Range:30
Rarity:Mythic
→
💰30,000
⭐Level 50+
Evolved Form

Rule (Room)

DPS:1100+189%
Range:45
Rarity:Mythical

📦 Required Materials

Surgical Tools×10
Heart Fragment×2

✨ New Ability

Gamma Knife

Internal destruction attack that stuns the target for 3 seconds and prevents health regeneration.

Cooldown: 35s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Shanks

DPS:480
Range:30
Rarity:Mythic
→
💰50,000
⭐Level 50+
Evolved Form

Shunks (Emperor)

DPS:1800+275%
Range:40
Rarity:Mythical

📦 Required Materials

Emperor's Haki×1
Griffin Shard×10

✨ New Ability

Divine Departure

A massive horizontal slash that knocks back and stuns all enemies hit.

Cooldown: 50s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Hatsune Miku

DPS:N/A
Range:25
Rarity:Mythic
→
💰20,000
⭐Level 50+
Evolved Form

Virtual Idon (Concert)

DPS:500N/A
Range:60
Rarity:Legendary

📦 Required Materials

Virtual Mic×5
Neon Crystal×20

✨ New Ability

World is Mine

Increases attack speed (SPA) of all units by 20% for 15 seconds.

Cooldown: 60s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Sasuke

DPS:470
Range:26
Rarity:Mythic
→
💰45,000
⭐Level 50+
Evolved Form

Sasuke (Eternal)

DPS:1700+262%
Range:38
Rarity:Mythical

📦 Required Materials

Eternal Eye×1
Dark Chakra×15

✨ New Ability

Indra's Arrow

Fires a lightning arrow that causes a massive explosion and paralyzes enemies.

Cooldown: 45s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Kirito

DPS:460
Range:18
Rarity:Mythic
→
💰25,000
⭐Level 50+
Evolved Form

Kriatu (Dual Wielder)

DPS:2100+357%
Range:20
Rarity:Mythical

📦 Required Materials

Dual Blade Fragment×10
Blue Crystal×5

✨ New Ability

Starburst Stream

A 16-hit combo that deals massive damage to a single target.

Cooldown: 30s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Sakuya Izayoi

DPS:490
Range:28
Rarity:Exclusive
→
💰45,000
⭐Level 50+
Evolved Form

Sakuya (Time Manipulator)

DPS:1200+145%
Range:50
Rarity:Mythical

📦 Required Materials

Silver Knife×20
Pocket Watch×1

✨ New Ability

Sakuya's World

Stops time for 7 seconds. Affected enemies take 20% more damage during the freeze.

Cooldown: 55s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Genos

DPS:450
Range:30
Rarity:Mythic
→
💰20,000
⭐Level 50+
Evolved Form

Cyborg Disciple (Core)

DPS:1300+189%
Range:45
Rarity:Legendary

📦 Required Materials

Cyborg Core×5
Metal Scrap×50

✨ New Ability

Incineration Cannon

Blasts a wide beam of energy that clears the entire path in front of him.

Cooldown: 40s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Akainu

DPS:510
Range:24
Rarity:Mythic
→
💰50,000
⭐Level 50+
Evolved Form

Fleet Admiral Magma

DPS:1650+224%
Range:42
Rarity:Mythical

📦 Required Materials

Magma Crystal×10
Admiral Cloak×1

✨ New Ability

Great Eruption

Massive magma fist strike that leaves a burning pool on the track.

Cooldown: 45s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Blackbeard

DPS:320
Range:25
Rarity:Legendary
→
💰40,000
⭐Level 50+
Evolved Form

Greybeard (Quake)

DPS:1550+384%
Range:35
Rarity:Mythical

📦 Required Materials

Quake Fruit Fragment×5
Dark Essence×10

✨ New Ability

Seaquake

Shakes the screen, dealing heavy damage and slowing all enemies by 50% for 5s.

Cooldown: 50s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Rukia

DPS:280
Range:20
Rarity:Legendary
→
💰35,000
⭐Level 50+
Evolved Form

Ruka (Bankai)

DPS:1400+400%
Range:32
Rarity:Mythical

📦 Required Materials

Snow Petal×15
Ice Soul×5

✨ New Ability

Hakka no Togame

Instant freeze on all enemies in range for 4 seconds with high shattering damage.

Cooldown: 45s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Gin Ichimaru

DPS:350
Range:35
Rarity:Legendary
→
💰35,000
⭐Level 50+
Evolved Form

Gen (Kamishini)

DPS:1600+357%
Range:70
Rarity:Mythical

📦 Required Materials

Snake Bone×10
Poison Extract×5

✨ New Ability

Buto Renjin

Rapid fire sword extensions that hit the same enemy 10 times.

Cooldown: 25s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Rock Lee

DPS:400
Range:12
Rarity:Legendary
→
💰15,000
⭐Level 50+
Evolved Form

Pebble (Eight Gates)

DPS:1800+350%
Range:15
Rarity:Legendary

📦 Required Materials

Weights of Effort×1
Training Seal×10

✨ New Ability

Night Moth

A high-speed kick that deals 5x damage but stuns Pebble for 5 seconds after use.

Cooldown: 30s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Shikamaru

DPS:290
Range:26
Rarity:Mythic
→
💰25,000
⭐Level 50+
Evolved Form

Shakumira (Shadow)

DPS:600+107%
Range:35
Rarity:Legendary

📦 Required Materials

Shadow Scroll×5
Ink Bottle×20

✨ New Ability

Shadow Strangle

Holds up to 10 enemies in place for 6 seconds.

Cooldown: 45s
View Base Unit View Evolved Unit →

Evolution Path

Base Form

Lelouch

DPS:350
Range:35
Rarity:Secret
→
💰75,000
⭐Level 50+
Evolved Form

Lulu (Emperor)

DPS:525+50%
Range:55
Rarity:Secret

📦 Required Materials

Geass Sigil×1
Chess Piece×10

✨ New Ability

Zero Requiem

All enemies in a massive radius are forced to walk back toward the entrance for 7 seconds. Additionally, affected enemies take 30% increased damage for the duration.

Cooldown: 45s
View Base Unit View Evolved Unit →

Ragna (The Silverware)

SecretDPS
⭐

Etherealization Stages

E2✓
Effect:

Extends transformation duration by 20 seconds

E4✓
Effect:

Shatter skill damage increased by 60% (reaching 285% total damage)

E6✓
Effect:

Unlocks core ability linking Fury stacks to damage

✨

Passive Abilities

Frost Born
Attacks freeze enemies. Frozen units suffer permanent 30% slow after thawing
💡

Strategy Notes

Ultimate burst unit with damage ceiling after transformation, but self-destructs. If you don't want Ragna to disappear after 2 minutes, you can intentionally avoid filling the energy bar.

🎯

Recommended Traits

Ruler

Jin Woo Monarch

SecretDPS
🔄

Base Evolution

Sun Jin Woo
→
Monarch
Materials Required:
Evolution Material IEvolution Material IIEvolution Material III
Evolution Changes:
  • Transforms from pure DPS to summoner
  • Base form can only attack ground, becomes Hybrid after upgrading to Mutilation
  • At max level deals ~32,000 damage with reduced attack frequency to 2.4s
⭐

Etherealization Stages

E2✓
Effect:

Increases shadow soldier count by 10

E4

No data available

E6✓
Effect:

Skill mana cost reduced by half

💡

Strategy Notes

Best character in the game. Summoner core, extremely fast at clearing Bosses, essential unit for Infinite and high-difficulty challenges.

🎯

Recommended Traits

Ruler Godly

Sasuke (Hebi)

MythicDPS
🔄

Base Evolution

Sasuke
→
Sasuke (Hebi)
Materials Required:
Cursed MarkCursed Sword
Evolution Changes:
  • Gains black flash mechanism (Amaterasu)
  • After evolution actually possesses Hybrid capability
  • Retains unit level but allows manual attribute point allocation
✨

Passive Abilities

Team 7 Synergy
When Kakashi or other Team 7 members are on field, gains 12% range and damage boost
Stun Chance
Base form has 50% chance to stun enemies, can be improved through etherealization after evolution
💡

Strategy Notes

Has unique evolution mechanism and black flash system, can handle flying units through Amaterasu passive after evolution. Recommend prioritizing SPA (attack speed) upgrades.

🎯

Recommended Traits

Ruler Sacred

Speedcart (Farm)

LegendarySupport
⭐

Etherealization Stages

E2

No data available

E4

No data available

E6✓
Effect:

Reaches maximum income

💡

Strategy Notes

Only gold production unit in the entire game, must etherealize to E6 for maximum income. Beginners must collect.

🎯

Recommended Traits

Superior

Evolution Strategy Tips

🎯

Prioritize DPS Units

Focus etherealization on your main damage dealers first. Units like Kenpachi, Sun Jin Woo, and Kyto benefit enormously from E6 upgrades.

💰

Farm Economic Units Early

Speedcart must reach E6 for maximum income efficiency. This should be one of your first etherealization priorities.

📊

Material Farming Routes

Legend Stages drop evolution materials. Use macros for efficient farming, and prioritize stages that drop materials for your key units.

⚡

Legendary vs Mythic Trade-off

Legendary units at E6 can compete with Mythic units at lower etherealization levels. Consider this when planning your resources.

Related Tools

You might also find these tools helpful

🔮

Evolution Calculator

Calculate total evolution costs and materials

→
📦

Materials Gallery

Browse evolution materials and farming guides

→
🗂️

Unit Database

Explore complete unit information and stats

→

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